Following the destruction of the Chimera's central tower in London and their armies eradicated throughout Britain in the Battle of London, humanity was aware that the Chimeran tower network was the key source of the Chimera's vulnerability. The Maquis, an European mainland resistance movement, approached the British for a joint-planned operation in destroying the Chimera's hub tower in Paris, France and liberating Europe as similar to the British's. However, the operation was not only to destroying the Paris central power, but also developing a virus created by Dr. Claude Bouchard that would eradicate the Chimera's Carrier strain that were tasked in transporting Chimeran virus-infected humans to conversion centers, and thus grounding the Chimera's conversion process to a halt.
Operation Overstrike is quite similar to Operation Deliverance in the sense that new troops are being transported to fight and liberate a country (in this case a continent) from the Chimera. The British Intelligence Corps would dispatch two squadrons of British forces to aid the Maquis.
The First Squadron under the command of Major Stephen Cartwright would fly first into Rotterdam, in which it is one of few open corridors to bypass the Chimera's fortifications, follow four hours later by the Second Squadron under Colonel Rachel Parker. Accompanied by the Maquis, the British would cross Chimera lines into Luxembourg where the Maquis stronghold lies, and then from there preparing their attack on the Paris Tower, hoping that its destruction would result in the meltdown of the tower network across Western Europe.
In the early phase of the operation, the Maquis made an major intelligence error in regarding to the Chimera's coastal defenses. Since the Chimera's occupation of Europe, the Chimera constructed massive fortifications the coastlines of Western Europe and making it almost impossible to penetrate through. The Chimera, however, hastily erected three gun towers in Rotterdam that came unnoticed for the Maquis and inflicting heavy casualties on Stephen Cartwright's First Squadron.
Despite the loss of human lives, human forces were able to disabled the gun towers and allowing Rachel Parker's Second Squadron to safely arrive, bringing supplies and reinforcements to a makeshift camp outside of Rotterdam. Stephen Cartwright would lead his task force to intercept Chimeran Earth Movers in Gerolstein that is heading for Luxembourg while Rachel Parker took the remainder of the Squadron to Luxembourg to finalize plans for the assault on the Paris hub tower.
During this time, British mercenary James Grayson, Colonel Roland Mallery, and Lieutenant Raine Bouchard investigated a new type of conversion facility in Bonn, Germany. The group is separated, with Bouchard captured by the Chimera and falling into the initial stages of the conversion process, but Grayson was able to rescue her, believing her to be yet unaffected, and then destroying the facility.
On September 2nd, Major Cartwright faced complications in stopping the Chimeran Earth Movers in Gerolstein and encroaching into Luxembourg. James Grayson aided Cartwright in immobilizing the Earth Movers were immobilized, but their victory is overshadowed by an all-out Chimeran assault on the fortress.
British and Maquis soldiers defend their base long enough to evacuate the personnel to their last stronghold in Reims, France. Dr. Claude Bouchard along with his daughter Raine were still occupied in developing the serum that would eradicate the Carrier strain, in which Raine had deliberately allowed herself to being infected in Bonn to allow her father to gain a sample of the Chimeran virus as to develop a serum. However, before the serum can be created, Dr. Bouchard is killed by the invading Chimera forces, but Raine was able to finish her father's work and gaining a sample. All surviving personnel and refugees fled to Reims, the only remaining safe zone left for the Maquis.
Following the evacuation to Reims, plans for the assault on the Paris tower were laid: Stephen Cartwright would lead the combined human military forces into battle. However, their goal is not only to destroy the hub tower, but also to distract the Chimera while allowing an infiltration into the Chrysalis Lair, a conversion facility where Chimera DNA are implanted into captured humans before they are taken to other conversion facilities, and releasing Dr. Bouchard's serum. Raine Bouchard volunteered for the infiltration, but was in fact already infected with the Chimeran virus and showing signs of falling to the Chimera's influence. James Grayson, having discovered Bouchard's infection, attempts to warn Mallery and the British about Bouchard, but only to be refuted by Mallery and imprisoned for treason.
Fortunately, Grayson was freed by Stephen Cartwright and Rachel Parker and given a second sample of the serum to take to the the Chrysalis Lair through the catacombs beneath Paris. At the same time Mallery's forces engaged the Chimera presence in the city and suffering a grave amount of human casualties. Grayson encountered Mallery who was infected with the Chimeran virus and purposefully preventing him from reaching the facility. Grayson killed Mallery and infiltrated the Chrysalis Lair, where he succeeded in using the serum just as Cartwright's forces destroyed the Paris tower. Just after Grayson used the serum, he discovered that Bouchard had mutated into a Chimeran entity calling itself as the Chrysalis and managed to kill it.
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The Operation Overstrike results in a major victory for humanity. Just as the tower fell in London, the main Chimeran tower was destroyed by Major Cartwright and his task force, presumably caused a chain reaction that destroys all the other towers along with the network and the entire Chimeran forces, everywhere in Western Europe had become disorganized, confused and begin to die. James Grayson also become a successful, he injected the Bouchard's serum into the "heart" of the Chrysalis Lair, infecting and killing all the Carrier strains and ground the conversion process to a halt Poland, Spain, the Balkans and Russia were still occupied by the Chimera, but after Overstrike these countries became the main battlefields within Northern, Eastern and Southern Europe, except for United Kingdom.
Despite eradication of the Carrier strain, however, the Chimera were able to overcome their setback by evolving and creating new strains among their ranks. Among them that succeed the Carriers was the Spinner strain. Furthermore, the Chimera finished the construction of a mass fleet of airships with their targets and destinations unknown at the time. The survived members of British and Maquis had made the memorial for those who died in the war in Europe.
- In Resistance: The Gathering Storm, Daedalus was looking through the eyes of other Chimeran species, in which he looked through the eyes of a Steelhead eating a human leg in Paris even though earlier Noah Grace was congratulating Stephen Cartwright for freeing western Europe. This can either be a mistake by the author or the Chimera trying to retake their lost territories like in Bracknell and Axbridge depicted in Resistance 2.